Dr John Scullion PhD

Dr John Scullion


Contact Details


Module Coordinator
Additional Lecturer


  • Disturbed/contaminated soils.
    • Carbon-nitrogen cycling and long-term effects of earthworm activity in former mine soils.
    • Impact of metals on microbial processes. Potential of phytoremediation for reducing soil metal and hydrocarbon concentrations.
    • Organic/ecological farming systems and soil processes.
    • Infectivity and efficiency of VA mycorrhizal fungi.
    • Earthworm-VA mycorrhizal fungi interactions. Earthworm population dynamics in conventional and organic arable rotations.
    • Soil ecology relationships to soil function. Impacts of climate change on soil ecology.
    • Chemometric and metabolomic approaches to the assessment of soil quality.
    • Biological treatment of wastewaters.



Al-Maliki, S.M.J., Jones, D., Godbold, D., Jones, D., Scullion, J. 2017. Elevated CO2 and Tree Species Affect Microbial Activity and Associated Aggregate Stability in Soil Amended with Litter. Forests 8 (3) 70 10.3390/f8030070 Cadair

Guo, Q., Meng, L., Humphreys, M.W., Scullion, J., Mur, L.A. 2017. Expression of FlHMA3, a vacuolar P1B2-ATPase from Festulolium loliaceum, correlates with response to cadmium stress. Plant Physiology and Biochemistry 112 pp. 270-277. 10.1016/j.plaphy.2017.01.013 Cadair


Crotty, F., Fychan, R., Sanderson, R., Rhymes, J., Bourdin, F., Scullion, J., Marley, C. 2016. Understanding the legacy effect of previous forage crop and tillage management on soil biology, after conversion to an arable crop rotation. Soil Biology and Biochemistry 103 pp. 241-252. 10.1016/j.soilbio.2016.08.018 Cadair

Humphreys, M., Loka, D., Gwynn-Jones, D., Scullion, J., Doonan, J., Gasior, D., Harper, J., Farrell, M., Kingston-Smith, A., Dodd, R., Chadwick, D., Hill, P., Mills, G., Hayes, F., Robinson, D., Jones, D. 2016. Climate-smart Strategies to Safeguard Persistency and Achieve Sustainable Grassland Production. 26th General Meeting of the EGF. Cadair

Hill, R., Saetnan, E.R., Scullion, J., Jones, D., Ostle, N.J., Edwards, A. 2016. Temporal and spatial influences incur reconfiguration of Arctic heathland soil bacterial community structure. Environmental Microbiology 18 (6) pp. 1942-1953. 10.1111/1462-2920.13017 Cadair

Lüscher, G., Ammari, Y., Andriets, A., Angelova, S., Arndorfer, M., Bailey, D., Balázs, K., Bogers, M., Bunce, R.G.H., Choisis, J., Dennis, P., Díaz, M., Dyman, T., Eiter, S., Fjellstad, W., Fraser, M., Friedel, J.K., Garchi, S., Geijzendorffer, I.R., Gomiero, T., González-bornay, G., Guteva, Y., Herzog, F., Jeanneret, P., Jongman, R.H.G., Kainz, M., Kwikiriza, N., López Díaz, M.L., Moreno, G., Nicholas-Davies, P., Nkwiine, C., Opio, J., Paoletti, M.G., Podmaniczky, L., Pointereau, P., Pulido, F., Sarthou, J., Schneider, M.K., Sghaier, T., Siebrecht, N., Stoyanova, S., Wolfrum, S., Yashchenko, S., Albrecht, H., Báldi, A., Belényesi, M., Benhadi-marin, J., Blick, T., Buholzer, S., Centeri, C., Choisis, N., Cuendet, G., De Lange, H.J., Déjean, S., Deltshev, C., Díaz Cosín, D.J., Dramstad, W., Elek, Z., Engan, G., Evtushenko, K., Falusi, E., Finch, O., Frank, T., Gavinelli, F., Genoud, D., Gillingham, P.K., Grónás, V., Gutiérrez, M., Häusler, W., Heer, X., Hübner, T., Isaia, M., Jerkovich, G., Jesus, J.B., Kakudidi, E., Kelemen, E., Koncz, N., Kovacs, E., Kovács-hostyánszki, A., Last, L., Ljubomirov, T., Mandery, K., Mayr, J., Mjelde, A., Muster, C., Nascimbene, J., Neumayer, J., Ødegaard, F., Ortiz Sánchez, F.J., Oschatz, M., Papaja-hülsbergen, S., Paschetta, M., Pavett, M., Pelosi, C., Penksza, K., Pommeresche, R., Popov, V., Radchenko, V., Richner, N., Riedel, S., Scullion, J., Sommaggio, D., Szalkovszki, O., Szerencsits, E., Trigo, D., Vale, J., Van Kats, R., Vasilev, A., Whittington, A.E., Wilkes-allemann, J., Zanetti, T. 2016. Farmland biodiversity and agricultural management on 237 farms in 13 European and two African regions. Ecology 97 (6) pp. 1625. 10.1890/15-1985.1 Other Cadair

Marshall, A.H., Collins, R.P., Humphreys, M.W., Scullion, J. 2016. A new emphasis on root traits for perennial grass and legume varieties with environmental and ecological benefits. Food and Energy Security 5 (1) pp. 26-39. 10.1002/fes3.78 Cadair

Jones, A.G., Bussell, J.S., Winters, A., Scullion, J., Jones, D. 2016. The functional quality of decomposing litter outputs from an Arctic plant community is affected by long-term exposure to enhanced UV-B. Ecological Indicators. Ecological Indicators 60 pp. 8-17. 10.1016/j.ecolind.2015.05.052 Cadair


Crotty, F., Fychan, R., Scullion, J., Sanderson, R., Marley, C. 2015. Assessing the impact of agricultural forage crops on soil biodiversity and abundance. Soil Biology and Biochemistry 91 pp. 119-126. 10.1016/j.soilbio.2015.08.036 Cadair

Jones, A.G., Scullion, J., Ostle, N.J., S, O., di Dio, A., Gwynn-Jones, D. 2015. Plant community composition and an insect outbreak influence phenol oxidase activity and soil-litter biochemistry in a sub-Arctic birch-heath. Polar Biology 38 (4) pp. 505-516. 10.1007/s00300-014-1613-8 Cadair

Crotty, F., Fychan, R., Scullion, J., Sanderson, R., Marley, C. 2015. Understanding the legacy effect of previous crop rotations on soil biology: Extended abstract. Improving Soil Health - Developing tools for land managers, Association of Applied Biologists. Cadair

Crotty, F., Fychan, R., Scullion, J., Sanderson, R., Marley, C. 2015. Linking legacy effects within crop rotations to the soil food web. Soil Ecology Society . Cadair

Fychan, R., Crotty, F., Scullion, J., Sanderson, R., Marley, C. 2015. Effects of forage species and sward-lifting on compacted soil. European Grassland Federation: Grassland and forages in high input dairy farms. Cadair

Detheridge, A., Scullion, J., Griffith, G., Marley, C.L., Fychan, R. 2015. A ground truthed DNA metabarcoding technique to determine fungal community structure in soils. Proceedings of AAB Conference : Healthy Soils - Developing Tools for Land Managers. Association of Applied Biologists Conference . Association of Applied Biologists Cadair

Gwynn-Jones, D., Jones, A., Ostle, N., Edwards, A., Scullion, J., Hill, R., Comont, D., Bussell, J., Oakley, S., Mason, K., Callaghan, T.V. 2015. How does increasing CO2 affect soil microbial diversity and carbon fluxes? In T. V. Callaghan., H. Savela. (eds) INTERACT Stories of Arctic Science. Aarhus University Press pp. 96-97. Other


Jones, A.G., Scullion, J., Ostle, N., Levy, P.E., Gwynn-Jones, D. 2014. Completing the FACE of elevated CO2 research. Environment International 73 pp. 252-258. 10.1016/j.envint.2014.07.021 Cadair

Crotty, F.V., Fychan, A.R., Scullion, J., Sanderson, R., Marley, C.L. 2014. The effects of agricultural forages on soil biology - linking the plant-soil-invertebrate ecosystem. In A. Hopkins. (ed) EGF at 50: The Future of European Grasslands: Grassland Science in Europe. 25th EGF Conference. pp. 267-269. Cadair

Scullion, J., Smith, A., Jones, D., Jones, D.L., Godbold, D.L. 2014. Deciduous woodland exposed to elevated atmospheric CO2 has species-specific impacts on anecic earthworms. Applied Soil Ecology 80 pp. 84-92. 10.1016/j.apsoil.2014.03.016 Cadair

Crotty, F., Fychan, R., Scullion, J., Sanderson, R., Marley, C. 2014. Investigating the impact of ecosystem change within agriculture on the soil food web. First Global Soil Biodiversity Initiative Conference. Cadair


Crotty, F.V., Fychan, A.R., Scullion, J., Davies, J.W., Scott, M.B., Sanderson, R., Marley, C.L. 2013. How agricultural management impacts the soil food web: comparing the effects of two different soil management treatments. Science and Practice for Grass-based Systems. 11th British Grassland Society Research Conference. Cadair

Crotty, F.V., Fychan, A.R., Scullion, J., Davies, J.W., Scott, M.B., Sanderson, R., Marley, C.L. 2013. How is the soil food web affected by management practices to promote high yielding and high quality grassland. Ento '13 International Symposium and Annual National Science Meeting.: Thirty years of Thornhill & Alcock: The Evolution of Insect Mating Systems. Ento '13 International Symposium and Annual National Science Meeting. Cadair

Crotty, F.V., Fychan, A.R., Scullion, J., Davies, J.W., Scott, M.B., Sanderson, R., Marley, C.L. 2013. Linking above ground management to below ground ecosystems: the effect of utilising alternative forages on the soil food web. Soil Ecology Society: Biodiversity and Function in Soil. 14th Biennial Soil Ecology Society Meeting 2013. Cadair

Humphreys, M., Farrell, M., Detheridge, A., Scullion, J., Kingston-Smith, A., O'Donovan, S.A. 2013. Resilient and multifunctional grasslands for agriculture and environmental service during a time of climate change. In Á. Helgadóttir., A. Hopkins. (eds) The Role of Grasslands in a Green Future: Threats and Perspectives in Less Favoured Areas. European Grassland Federation pp. 335-337. Cadair

Al-Maliki, S., Scullion, J. 2013. Interactions between earthworms and residues of differing quality affecting aggregate stability and microbial dynamics. Applied Soil Ecology 64 pp. 56-62. 10.1016/j.apsoil.2012.10.008 Other Cadair


Gwynn-Jones, D., Jones, A.G., Waterhouse, A., Winters, A., Comont, D., Scullion, J., Gardias, R., Graee, B.J., Lee, J.A., Callaghan, T.V. 2012. Enhanced UV-B and Elevated CO2 Impacts Sub-Arctic Shrub Berry Abundance, Quality and Seed Germination. Ambio: A Journal of the Human Environment 41 (3) pp. 256-268. 10.1007/s13280-012-0311-4 Cadair

Bussell, J.S., Gwynn-Jones, D., Griffith, G.W., Scullion, J. 2012. Above- and below-ground responses of Calamagrostis purpurea to UV-B radiation and elevated CO2 under phosphorus limitation. Physiologia Plantarum 145 (4) pp. 619-628. 10.1111/j.1399-3054.2012.01595.x Cadair


Marley, C.L., Powell, H.G., Scullion, J., Griffith, G.W., Fychan, R., McCalman, H.M.Smith, S.P. 2011. The PROSOIL project - Promoting healthy soils in Wales. Other Cadair

Jones, A., Scullion, J., Gwynn-Jones, D.Smith, S.P., Spikes, K.J. 2011. Soils as carbon sinks or sources: From mid-Wales to the Arctic. Other Cadair


Matthews, R., Winson, M.K., Scullion, J. 2009. Treating landfill leachate using passive aeration trickling filters; effects of leachate characteristics and temperature on rates and process dynamics. Science of the Total Environment 407 (8) pp. 2557-2564. 10.1016/j.scitotenv.2009.01.034 Cadair


Gee, J.H.R., Barber, I., Scullion, J. 2008. Baseline Report for the Afon Teifi Restoration Project. CCW Contract Science Report CCW/WW/08/6 Cyngor Cefn Gwlad Cymru Cadair


Elliott, G.N., Worgan, H., Broadhurst, D.I., Draper, J.H., Scullion, J. 2007. Soil differentiation using fingerprint Fourier transform infrared spectroscopy, chemometrics and genetic algorithm-based feature selection. Soil Biology and Biochemistry 39 (11) pp. 2888-2896. 10.1016/j.soilbio.2007.05.032 Cadair

Scullion, J., Neal, S., Philips, L. 2007. Earthworm casting and burrowing activity in conventional and organic grass-arable rotations. European Journal of Soil Biology 43 (1) pp. S216-S221. 10.1016/j.ejsobi.2007.08.018 Cadair

Scullion, J., Winson, M.K., Matthews, R. 2007. Inhibition and recovery in a fixed microbial film leachate treatment system subject to shock loading of copper and zinc. Water Research 41 (18) pp. 4129-4138. 10.1016/j.watres.2007.05.043 Cadair


Baker, G.H., Brown, G., Butt, K., Curry, J.P., Scullion, J. 2006. Introduced earthworms in agricultural and reclaimed land: their ecology and influences on soil properties, plant production and other soil biota. In P. F. Hendrix. (ed) Biological Invasions Belowground: Earthworms as Invasive Species. 2006th edn, Springer Nature pp. 101-116. 10.1007/978-1-4020-5429-7_10 Cadair

Scullion, J. 2006. Remediating polluted soils. The Science of Nature Naturwissenschaften 93 (2) pp. 51-65. 10.1007/s00114-005-0079-5 Cadair


Huang, W., Goodacre, R., Elliott, G.N., Beckmann, M., Worgan, H., Bailey, M.J., Williams, P.A., Scullion, J., Draper, J. 2005. The use of chemical profiling for monitoring metabolic changes in artificial soil slurries caused by horizontal gene transfer. Metabolomics pp. 305-315. 10.1007/s11306-005-0011-1 Cadair


Marashi, A.R.A., Scullion, J. 2004. Porosity and hydrological changes in surface mine soils. ISCO 2004 - 13th International Soil Conservation Organisation Conference. Cadair Other


Marashi, A.R.A., Scullion, J. 2003. Earthworm casts form stable aggregates in physically degraded soils. Biology and Fertility of Soils 37 (6) pp. 375-380. 10.1007/s00374-003-0617-2 Cadair

Scullion, J., Goodacre, R., Elliott, G.N., Huang, W., Worgan, H., Gwynn-Jones, D., Griffith, G.W., Darby, R.M., Bailey, M.J., Clegg, C.D., Draper, J.H. 2003. Food quality and microbial succession in ageing earthworm casts: standard microbial indices and metabolic fingerprinting. 10.1078/0031-4056-00276 Cadair

Gwynn-Jones, D., Huang, W., Easton, G.L., Goodacre, R., Scullion, J. 2003. UV-B radiation induced changes in litter quality affects earthworm growth and cast characteristics as determined by metabolic fingerprinting. Pedobiologia 47 (5-6) pp. 784-787. 10.1078/0031-4056-00258 Cadair

Scullion, J., Elliott, G.N., Huang, W., Goodacre, R., Worgan, H., Darby, R.M., Bailey, M.J., Gwynn-Jones, D., Griffith, G.W., Winson, M.K., Williams, P.A., Clegg, C.D., Draper, J. 2003. Use of earthworm casts to validate FT-IR spectroscopy as a 'sentinel' technology for high-throughput monitoring of global changes in microbial ecology. Pedobiologia 47 (5-6) 7th International Symposium on Earthworm Ecology. pp. 440-446. 10.1078/0031-4056-00210 Cadair


Scullion, J., Neale, S., Philipps, L. 2002. Comparisons of earthworm populations and cast properties in conventional and organic arable rotations. Soil Use and Management 18 (S1) pp. 293-300. 10.1111/j.1475-2743.2002.tb00271.x Cadair

Tuffen, F., Eason, W.R., Scullion, J. 2002. The effect of earthworms and arbuscular mycorrhizal fungi on growth of and 32P transfer between Allium porrum plants. Soil Biology and Biochemistry 34 (7) pp. 1027-1036. 10.1016/S0038-0717(02)00036-6 Cadair

Khan, M., Scullion, J. 2002. Effects of metal (Cd, Cu, Ni, Pb or Zn) enrichment of sewage-sludge on soil micro-organisms and their activities. Applied Soil Ecology 20 (2) pp. 145-155. 10.1016/S0929-1393(02)00018-5 Cadair

Oudeh, M., Khan, M., Scullion, J. 2002. Plant accumulation of potentially toxic elements in sewage sludge as affected by soil organic matter level and mycorrhizal fungi. Environmental Pollution 116 (2) pp. 293-300. 10.1016/S0269-7491(01)00128-2 Cadair

Siddiqui, S., Adams, W.A., Scullion, J. 2002. The phytotoxicity and degradation of diesel hydrocarbons in soil. Journal of Plant Nutrition and Soil Science 164 (6) pp. 631-635. 10.1002/1522-2624(200112)164:6<631::AID-JPLN631>3.0.CO;2-E Cadair


Scullion, J., Malik, A. 2000. Earthworm activity affecting organic matter, aggregation and microbial activity in soils restored after opencast mining for coal. Soil Biology and Biochemistry 32 (1) pp. 119-126. 10.1016/S0038-0717(99)00142-X Cadair