Dr John Scullion


Dr John Scullion


Contact Details


Module Coordinator
Additional Lecturer


  • Disturbed/contaminated soils.
    • Carbon-nitrogen cycling and long-term effects of earthworm activity in former mine soils.
    • Impact of metals on microbial processes. Potential of phytoremediation for reducing soil metal and hydrocarbon concentrations.
    • Organic/ecological farming systems and soil processes.
    • Infectivity and efficiency of VA mycorrhizal fungi.
    • Earthworm-VA mycorrhizal fungi interactions. Earthworm population dynamics in conventional and organic arable rotations.
    • Soil ecology relationships to soil function. Impacts of climate change on soil ecology.
    • Chemometric and metabolomic approaches to the assessment of soil quality.
    • Biological treatment of wastewaters.


Vegetation and edaphic factors influence rapid establishment of distinct fungal communities on former coal-spoil sitesDetheridge, A., Comont, D. C., Callaghan, A., Bussell, J. S., Brand, G., Jones, D., Scullion, J. & Griffith, G. 2018 In : Fungal Ecology.p. 92-103
Elevated CO2 and Tree Species Affect Microbial Activity and Associated Aggregate Stability in Soil Amended with LitterAl-Maliki, S. M. J., Jones, D., Godbold, D., Jones, D. & Scullion, J. 2017 In : Forests.8, 3, 70
Expression of FlHMA3, a vacuolar P1B2-ATPase from Festulolium loliaceum, correlates with response to cadmium stressGuo, Q., Meng, L., Humphreys, M. W., Scullion, J. & Mur, L. A. J. 2017 In : Plant Physiology and Biochemistry.112, p. 270-2778 p.
Understanding the legacy effect of previous forage crop and tillage management on soil biology, after conversion to an arable crop rotationCrotty, F., Fychan, R., Sanderson, R., Rhymes, J., Bourdin, F., Scullion, J. & Marley, C. 2016 In : Soil Biology and Biochemistry.103, p. 241-252
Climate-smart Strategies to Safeguard Persistency and Achieve Sustainable Grassland ProductionHumphreys, M., Loka, D., Gwynn-Jones, D., Scullion, J., Doonan, J., Gasior, D., Harper, J., Farrell, M., Kingston-Smith, A., Dodd, R., Chadwick, D., Hill, P., Mills, G., Hayes, F., Robinson, D. & Jones, D. 2016
Temporal and spatial influences incur reconfiguration of Arctic heathland soil bacterial community structureHill, R., Saetnan, E. R., Scullion, J., Jones, D., Ostle, . N. J. & Edwards, A. 2016 In : Environmental Microbiology.18, 6, p. 1942-195312 p.
Farmland biodiversity and agricultural management on 237 farms in 13 European and two African regionsLüscher, G., Ammari, Y., Andriets, A., Angelova, S., Arndorfer, M., Bailey, D., Balázs, K., Bogers, M., Bunce, R. G. H., Choisis, J., Dennis, P., Díaz, M., Dyman, T., Eiter, S., Fjellstad, W., Fraser, M., Friedel, J. K., Garchi, S., Geijzendorffer, I. R., Gomiero, T., González-bornay, G., Guteva, Y., Herzog, F., Jeanneret, P., Jongman, R. H. G., Kainz, M., Kwikiriza, N., López Díaz, M. L., Moreno, G., Nicholas-Davies, P., Nkwiine, C., Opio, J., Paoletti, M. G., Podmaniczky, L., Pointereau, P., Pulido, F., Sarthou, J., Schneider, M. K., Sghaier, T., Siebrecht, N., Stoyanova, S., Wolfrum, S., Yashchenko, S., Albrecht, H., Báldi, A., Belényesi, M., Benhadi-marin, J., Blick, T., Buholzer, S., Centeri, C., Choisis, N., Cuendet, G., De Lange, H. J., Déjean, S., Deltshev, C., Díaz Cosín, D. J., Dramstad, W., Elek, Z., Engan, G., Evtushenko, K., Falusi, E., Finch, O., Frank, T., Gavinelli, F., Genoud, D., Gillingham, P. K., Grónás, V., Gutiérrez, M., Häusler, W., Heer, X., Hübner, T., Isaia, M., Jerkovich, G., Jesus, J. B., Kakudidi, E., Kelemen, E., Koncz, N., Kovacs, E., Kovács-hostyánszki, A., Last, L., Ljubomirov, T., Mandery, K., Mayr, J., Mjelde, A., Muster, C., Nascimbene, J., Neumayer, J., Ødegaard, F., Ortiz Sánchez, F. J., Oschatz, M., Papaja-hülsbergen, S., Paschetta, M., Pavett, M., Pelosi, C., Penksza, K., Pommeresche, R., Popov, V., Radchenko, V., Richner, N., Riedel, S., Scullion, J., Sommaggio, D., Szalkovszki, O., Szerencsits, E., Trigo, D., Vale, J., Van Kats, R., Vasilev, A., Whittington, A. E., Wilkes-allemann, J. & Zanetti, T. 2016 In : Ecology.97, 6, p. 1625
A new emphasis on root traits for perennial grass and legume varieties with environmental and ecological benefitsMarshall, A. H., Collins, R. P., Humphreys, M. W. & Scullion, J. 2016 In : Food and Energy Security.5, 1, p. 26-3914 p.
The functional quality of decomposing litter outputs from an Arctic plant community is affected by long-term exposure to enhanced UV-B. Ecological Indicators.Jones, A. G., Bussell, J. S., Winters, A., Scullion, J. & Jones, D. 2016 In : Ecological Indicators.60, p. 8-1710 p.
Assessing the impact of agricultural forage crops on soil biodiversity and abundanceCrotty, F., Fychan, R., Scullion, J., Sanderson, R. & Marley, C. 2015 In : Soil Biology and Biochemistry.91, p. 119-126
Plant community composition and an insect outbreak influence phenol oxidase activity and soil-litter biochemistry in a sub-Arctic birch-heathJones, A. G., Scullion, J., Ostle, N. J., S, O., di Dio, A. & Gwynn-Jones, D. 2015 In : Polar Biology.38, 4, p. 505-516
Understanding the legacy effect of previous crop rotations on soil biology: Extended abstractCrotty, F., Fychan, R., Scullion, J., Sanderson, R. & Marley, C. 2015
Linking legacy effects within crop rotations to the soil food webCrotty, F., Fychan, R., Scullion, J., Sanderson, R. & Marley, C. 2015
Effects of forage species and sward-lifting on compacted soilFychan, R., Crotty, F., Scullion, J., Sanderson, R. & Marley, C. 2015
A ground truthed DNA metabarcoding technique to determine fungal community structure in soilsDetheridge, A., Scullion, J., Griffith, G., Marley, C. L. & Fychan, R. 2015 Proceedings of AAB Conference : Healthy Soils - Developing Tools for Land Managers.Association of Applied Biologists
How does increasing CO2 affect soil microbial diversity and carbon fluxes?Gwynn-Jones, D., Jones, A., Ostle, . N., Edwards, A., Scullion, J., Hill, R., Comont, D., Bussell, J., Oakley, S., Mason, K. & Callaghan, T. V. 2015 INTERACT Stories of Arctic Science. Callaghan, T. V. & Savela, H. (eds.). Aarhus University Press, p. 96-972 p.
Completing the FACE of elevated CO2 researchJones, A. G., Scullion, J., Ostle, N., Levy, P. E. & Gwynn-Jones, D. 2014 In : Environment International.73, p. 252-2587 p.
The effects of agricultural forages on soil biology - linking the plant-soil-invertebrate ecosystemCrotty, F. V., Fychan, A. R., Scullion, J., Sanderson, R. & Marley, C. L. 2014 EGF at 50: The Future of European Grasslands: Grassland Science in Europe. Hopkins, A. (ed.). Vol. 19, p. 267-2693 p.
Deciduous woodland exposed to elevated atmospheric CO2 has species-specific impacts on anecic earthwormsScullion, J., Smith, A., Jones, D., Jones, D. L. & Godbold, D. L. 2014 In : Applied Soil Ecology.80, p. 84-929 p.
Investigating the impact of ecosystem change within agriculture on the soil food web.Crotty, F., Fychan, R., Scullion, J., Sanderson, R. & Marley, C. 2014
Interactions between earthworms and residues of differing quality affecting aggregate stability and microbial dynamicsAl-Maliki, S. & Scullion, J. 2013 In : Applied Soil Ecology.64, p. 56-627 p.
How agricultural management impacts the soil food web: comparing the effects of two different soil management treatments.Crotty, F. V., Fychan, A. R., Scullion, J., Davies, J. W., Scott, M. B., Sanderson, R. & Marley, C. L. 2013 Science and Practice for Grass-based Systems.
How is the soil food web affected by management practices to promote high yielding and high quality grasslandCrotty, F. V., Fychan, A. R., Scullion, J., Davies, J. W., Scott, M. B., Sanderson, R. & Marley, C. L. 2013 Ento '13 International Symposium and Annual National Science Meeting.: Thirty years of Thornhill & Alcock: The Evolution of Insect Mating Systems.
Linking above ground management to below ground ecosystems: the effect of utilising alternative forages on the soil food web.Crotty, F. V., Fychan, A. R., Scullion, J., Davies, J. W., Scott, M. B., Sanderson, R. & Marley, C. L. 2013 Soil Ecology Society: Biodiversity and Function in Soil.
Resilient and multifunctional grasslands for agriculture and environmental service during a time of climate changeHumphreys, M., Farrell, M., Detheridge, A., Scullion, J., Kingston-Smith, A. & O'Donovan, S. A. 2013 The Role of Grasslands in a Green Future: Threats and Perspectives in Less Favoured Areas. Helgadóttir, Á. & Hopkins, A. (eds.). European Grassland Federation, p. 335-3373 p.
Enhanced UV-B and Elevated CO2 Impacts Sub-Arctic Shrub Berry Abundance, Quality and Seed GerminationGwynn-Jones, D., Jones, A. G., Waterhouse, A., Winters, A., Comont, D., Scullion, J., Gardias, R., Graee, B. J., Lee, J. A. & Callaghan, T. V. 2012 In : Ambio: A Journal of the Human Environment.41, 3, p. 256-26813 p.
Above- and below-ground responses of Calamagrostis purpurea to UV-B radiation and elevated CO2 under phosphorus limitationBussell, J. S., Gwynn-Jones, D., Griffith, G. W. & Scullion, J. 2012 In : Physiologia Plantarum.145, 4, p. 619-62810 p.
The PROSOIL project - Promoting healthy soils in WalesMarley, C. L., Powell, H. G., Scullion, J., Griffith, G. W., Fychan, R., McCalman, H. M. & Smith, S. P. (ed.) 2011 IBERS Knowledge-based Innovations, 4, p. 5-95 p.
Soils as carbon sinks or sources: From mid-Wales to the ArcticJones, A., Scullion, J., Gwynn-Jones, D., Smith, S. P. (ed.) & Spikes, K. J. (ed.) 2011 IBERS Knowledge-based Innovations, 4, p. 11-155 p.
Treating landfill leachate using passive aeration trickling filters; effects of leachate characteristics and temperature on rates and process dynamicsMatthews, R., Winson, M. K. & Scullion, J. 2009 In : Science of the Total Environment.407, 8, p. 2557-25648 p.
Baseline Report for the Afon Teifi Restoration ProjectGee, J. H. R., Barber, I. & Scullion, J. 2008 Cyngor Cefn Gwlad Cymru. 73 p. (CCW Contract Science Report CCW/WW/08/6)
Soil differentiation using fingerprint Fourier transform infrared spectroscopy, chemometrics and genetic algorithm-based feature selectionElliott, G. N., Worgan, H., Broadhurst, D. I., Draper, J. H. & Scullion, J. 2007 In : Soil Biology and Biochemistry.39, 11, p. 2888-28969 p.
Earthworm casting and burrowing activity in conventional and organic grass-arable rotationsScullion, J., Neal, S. & Philips, L. 2007 In : European Journal of Soil Biology.43, 1, p. S216-S221
Inhibition and recovery in a fixed microbial film leachate treatment system subject to shock loading of copper and zincScullion, J., Winson, M. K. & Matthews, R. 2007 In : Water Research.41, 18, p. 4129-413810 p.
Introduced earthworms in agricultural and reclaimed land: their ecology and influences on soil properties, plant production and other soil biotaBaker, G. H., Brown, G., Butt, K., Curry, J. P. & Scullion, J. 2006 Biological Invasions Belowground: Earthworms as Invasive Species. Hendrix, P. F. (ed.). 2006 ed.Springer Nature, p. 101-11616 p.
Remediating polluted soilsScullion, J. 2006 In : Naturwissenschaften.93, 2, p. 51-6515 p.
The use of chemical profiling for monitoring metabolic changes in artificial soil slurries caused by horizontal gene transferHuang, W., Goodacre, R., Elliott, G. N., Beckmann, M., Worgan, H., Bailey, M. J., Williams, P. A., Scullion, J. & Draper, J. 2005 In : Metabolomics.p. 305-31511 p.
Porosity and hydrological changes in surface mine soilsMarashi, A. R. A. & Scullion, J. 2004
Earthworm casts form stable aggregates in physically degraded soilsMarashi, A. R. A. & Scullion, J. 2003 In : Biology and Fertility of Soils.37, 6, p. 375-3806 p.
Food quality and microbial succession in ageing earthworm casts: standard microbial indices and metabolic fingerprintingScullion, J., Goodacre, R., Elliott, G. N., Huang, W., Worgan, H., Gwynn-Jones, D., Griffith, G. W., Darby, R. M., Bailey, M. J., Clegg, C. D. & Draper, J. H. 2003 p. 5-62 p.
UV-B radiation induced changes in litter quality affects earthworm growth and cast characteristics as determined by metabolic fingerprintingGwynn-Jones, D., Huang, W., Easton, G. L., Goodacre, R. & Scullion, J. 2003 In : Pedobiologia.47, 5-6, p. 784-7872 p.
Use of earthworm casts to validate FT-IR spectroscopy as a 'sentinel' technology for high-throughput monitoring of global changes in microbial ecologyScullion, J., Elliott, G. N., Huang, W., Goodacre, R., Worgan, H., Darby, R. M., Bailey, M. J., Gwynn-Jones, D., Griffith, G. W., Winson, M. K., Williams, P. A., Clegg, C. D. & Draper, J. 2003 In : Pedobiologia.47, 5-6, p. 440-4467 p.
Comparisons of earthworm populations and cast properties in conventional and organic arable rotationsScullion, J., Neale, S. & Philipps, L. 2002 In : Soil Use and Management.18, S1, p. 293-3008 p.
The effect of earthworms and arbuscular mycorrhizal fungi on growth of and 32P transfer between Allium porrum plantsTuffen, F., Eason, W. R. & Scullion, J. 2002 In : Soil Biology and Biochemistry.34, 7, p. 1027-103610 p.
Effects of metal (Cd, Cu, Ni, Pb or Zn) enrichment of sewage-sludge on soil micro-organisms and their activitiesKhan, M. & Scullion, J. 2002 In : Applied Soil Ecology.20, 2, p. 145-15511 p.
Plant accumulation of potentially toxic elements in sewage sludge as affected by soil organic matter level and mycorrhizal fungiOudeh, M., Khan, M. & Scullion, J. 2002 In : Environmental Pollution.116, 2, p. 293-3008 p.
The phytotoxicity and degradation of diesel hydrocarbons in soilSiddiqui, S., Adams, W. A. & Scullion, J. 2002 In : Journal of Plant Nutrition and Soil Science.164, 6, p. 631-6355 p.
Earthworm activity affecting organic matter, aggregation and microbial activity in soils restored after opencast mining for coalScullion, J. & Malik, A. 2000 In : Soil Biology and Biochemistry.32, 1, p. 119-1268 p.